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Larger ear surfaces contain more capillaries, and more heat can be released. Of all the elephants, African bush elephants live in the hottest climates, and have the largest ear flaps.
The trunk, or proboscis , is a fusion of the nose and upper lip, although in early fetal life, the upper lip and trunk are separated. It contains up to , separate muscle fascicles , with no bone and little fat.
These paired muscles consist of two major types: superficial surface and internal. The former are divided into dorsals, ventrals , and laterals while the latter are divided into transverse and radiating muscles.
The muscles of the trunk connect to a bony opening in the skull. The nasal septum is composed of tiny muscle units that stretch horizontally between the nostrils.
Cartilage divides the nostrils at the base. The muscles work both with and against each other. A unique proboscis nerve — formed by the maxillary and facial nerves — runs along both sides of the trunk.
Elephant trunks have multiple functions, including breathing, olfaction , touching, grasping, and sound production. The African elephant has two finger-like extensions at the tip of the trunk that allow it to grasp and bring food to its mouth.
The Asian elephant has only one, and relies more on wrapping around a food item and squeezing it into its mouth.
One elephant has been observed to graze by kneeling on its front legs, raising on its hind legs and taking in grass with its lips.
Elephants usually have 26 teeth: the incisors , known as the tusks , 12 deciduous premolars , and 12 molars. Unlike most mammals, which grow baby teeth and then replace them with a single permanent set of adult teeth, elephants are polyphyodonts that have cycles of tooth rotation throughout their lives.
The chewing teeth are replaced six times in a typical elephant's lifetime. Teeth are not replaced by new ones emerging from the jaws vertically as in most mammals.
Instead, new teeth grow in at the back of the mouth and move forward to push out the old ones. The first chewing tooth on each side of the jaw falls out when the elephant is two to three years old.
The second set of chewing teeth falls out at four to six years old. The third set falls out at 9—15 years of age, and set four lasts until 18—28 years of age.
The fifth set of teeth falls out at the early 40s. The sixth and usually final set must last the elephant the rest of its life.
Elephant teeth have loop-shaped dental ridges, which are thicker and more diamond-shaped in African elephants. The tusks of an elephant are modified second incisors in the upper jaw.
A newly developed tusk has a smooth enamel cap that eventually wears off. The dentine is known as ivory and its cross-section consists of crisscrossing line patterns, known as "engine turning", which create diamond-shaped areas.
As a piece of living tissue, a tusk is relatively soft; it is as hard as the mineral calcite. Much of the tusk can be seen outside; the rest is in a socket in the skull.
At least one-third of the tusk contains the pulp and some have nerves stretching to the tip. Thus it would be difficult to remove it without harming the animal.
When removed, ivory begins to dry up and crack if not kept cool and moist. Tusks serve multiple purposes. They are used for digging for water, salt, and roots; debarking or marking trees; and for moving trees and branches when clearing a path.
When fighting, they are used to attack and defend, and to protect the trunk. Like humans, who are typically right- or left-handed , elephants are usually right- or left-tusked.
The dominant tusk, called the master tusk, is generally more worn down, as it is shorter with a rounder tip. In the Asian species, only the males have large tusks.
Female Asians have very small tusks, or none at all. Hunting for elephant ivory in Africa  and Asia  has led to natural selection for shorter tusks   and tusklessness.
An elephant's skin is generally very tough, at 2. The skin around the mouth, anus , and inside of the ear is considerably thinner. Elephants typically have grey skin, but African elephants look brown or reddish after wallowing in coloured mud.
Asian elephants have some patches of depigmentation, particularly on the forehead and ears and the areas around them. Calves have brownish or reddish hair, especially on the head and back.
As elephants mature, their hair darkens and becomes sparser, but dense concentrations of hair and bristles remain on the end of the tail as well as the chin, genitals and the areas around the eyes and ear openings.
Normally the skin of an Asian elephant is covered with more hair than its African counterpart. An elephant uses mud as a sunscreen, protecting its skin from ultraviolet light.
Although tough, an elephant's skin is very sensitive. Without regular mud baths to protect it from burning, insect bites and moisture loss, an elephant's skin suffers serious damage.
After bathing, the elephant will usually use its trunk to blow dust onto its body and this dries into a protective crust.
Elephants have difficulty releasing heat through the skin because of their low surface-area-to-volume ratio , which is many times smaller than that of a human.
They have even been observed lifting up their legs, presumably in an effort to expose their soles to the air. To support the animal's weight, an elephant's limbs are positioned more vertically under the body than in most other mammals.
The long bones of the limbs have cancellous bone in place of medullary cavities. This strengthens the bones while still allowing haematopoiesis.
Elephants are incapable of rotating their front legs, as the ulna and radius are fixed in pronation ; the "palm" of the manus faces backward.
Elephants can move both forwards and backwards, but cannot trot , jump , or gallop. They use only two gaits when moving on land: the walk and a faster gait similar to running.
With no "aerial phase", the fast gait does not meet all the criteria of running, although the elephant uses its legs much like other running animals, with the hips and shoulders falling and then rising while the feet are on the ground.
Spring-like kinetics could explain the difference between the motion of elephants and other animals. The brain of an elephant weighs 4.
While the elephant brain is larger overall, it is proportionally smaller. The cerebrum and cerebellum are well developed, and the temporal lobes are so large that they bulge out laterally.
The vocal folds are long and are attached close to the epiglottis base. When comparing an elephant's vocal folds to those of a human, an elephant's are longer, thicker, and have a larger cross-sectional area.
In addition, they are tilted at 45 degrees and positioned more anteriorly than a human's vocal folds. It has a double-pointed apex , an unusual trait among mammals.
Unlike many other animals, the heart rate speeds up by 8 to 10 beats per minute when the elephant is lying down. This may allow the animal to deal with the pressure differences when its body is underwater and its trunk is breaking the surface for air,  although this explanation has been questioned.
The majority of an elephant's food intake goes undigested despite the process lasting up to a day. A male elephant's testes are located internally near the kidneys.
It is S-shaped when fully erect and has a Y-shaped orifice. The vulva is located between the hind legs instead of near the tail as in most mammals.
Determining pregnancy status can be difficult due to the animal's large abdominal cavity. The female's mammary glands occupy the space between the front legs, which puts the suckling calf within reach of the female's trunk.
The core body temperature averages The African bush elephant can be found in habitats as diverse as dry savannahs , deserts , marshes , and lake shores, and in elevations from sea level to mountain areas above the snow line.
Forest elephants mainly live in equatorial forests but will enter gallery forests and ecotones between forests and savannahs.
Elephants tend to stay near water sources. At midday, elephants rest under trees and may doze off while standing. Sleeping occurs at night while the animal is lying down.
Elephants go on seasonal migrations in search of food, water, minerals, and mates. Because of their large size, elephants have a huge impact on their environments and are considered keystone species.
Their habit of uprooting trees and undergrowth can transform savannah into grasslands; when they dig for water during drought, they create waterholes that can be used by other animals.
They can enlarge waterholes when they bathe and wallow in them. At Mount Elgon , elephants excavate caves that are used by ungulates , hyraxes, bats, birds and insects.
The seeds are typically dispersed in large amounts over great distances. This ecological niche cannot be filled by the next largest herbivore, the tapir.
At Murchison Falls National Park in Uganda, the overabundance of elephants has threatened several species of small birds that depend on woodlands.
Their weight can compact the soil, which causes the rain to run off , leading to erosion. Elephants typically coexist peacefully with other herbivores, which will usually stay out of their way.
Some aggressive interactions between elephants and rhinoceros have been recorded. At Aberdare National Park , Kenya, a rhino attacked an elephant calf and was killed by the other elephants in the group.
This is due to lower predation pressures that would otherwise kill off many of the individuals with significant parasite loads. Female elephants spend their entire lives in tight-knit matrilineal family groups, some of which are made up of more than ten members, including three mothers and their dependent offspring, and are led by the matriarch which is often the eldest female.
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